Table 3.

Commonly used immunophenotypic and genetic markers in the diagnosis of NK- and T-cell neoplasms

MarkerDiagnostic utilityInterpretation and comments
Antigens assessed by immunohistochemistry (IHC) or flow cytometry (FC) 
Pan-T-cell antigens:
CD2, CD3, CD5, CD7, CD43 (IHC, FC) 
Identification T-cell (or NK) populations.
Aberrant loss of T-cell antigen expression is a phenotypic marker favoring neoplastic over benign lymphoproliferation. 
NK and T cells are distinguished by the expression of surface CD3 (assessed on fresh cells by FC), positive in T cells and negative in NK cells, whereas both T and NK cells express cytoplasmic CD3 (assessed by IHC on routinely processed tissues); in addition, CD5 is constitutively absent in NK cells (and in TCRγδ+ T cells)
Extensive loss of T-cell antigens and possibly «null» immunophenotype is characteristic of ALK+ and to a lesser extent ALK ALCL.
CD7 loss is typical of ATLL. 
CD4 and CD8 (IHC, FC) Characterization of T-cell proliferations according to the CD4+CD8 or CD4CD8+ lineages. Double negative expression may indicate immature lymphoid cells, TCRγδ+ or NK cells, or aberrant phenotype in mature TCRαβ+ T cells, found in ALPS or lymphomas.
Double positive expression may indicate immature lymphoid cells, or aberrant phenotype in mature TCRαβ+ T cells, usually indicative of malignancy. 
TCRβ and TCRδ chains (IHC, FC) Identification of TCRαβ+ vs TCRγδ+ T cells. Most nodal PTCLs derive from TCRαβ+ T cells.
PTCL may show TCR downregulation, or silencing of TCR expression. 
TRBC1 (IHC, FC) Identification of clonal T-cell population by monotypic expression pattern. Only applicable if surface CD3 and TCRαβ expression is retained. 
CD30 (ICH, FC) Activation marker, not lineage-specific.
Strong uniform expression in ALCLs, heterogeneous expression in a proportion of cases in many TCL entities, usually in a subset of the cells. 
Only scattered cells may be positive in small-cell and histiocyte-rich patterns of ALK+ ALCL.
CD30 expression in bystander Reed–Sternberg-like cells in TFHL and PTCL-NOS.
May be targetable by antibody–drug conjugates. 
CD15 (IHC, FC) Glycan determinant, marker of myeloid cells.
May be positive in the neoplastic cells in ALCL and a subset of PTCL-NOS. 
Coexpression of CD30 and CD15 in bystander HRS B cells in various PTCLs. 
ALK (IHC) Demonstration of ALK expression is essential for the diagnosis of ALK+ ALCL. IHC is an excellent surrogate to ALK gene fusion detection by genetic assays. Variable patterns of subcellular staining according to the partner gene of ALK in the fusion.
Nuclear and cytoplasmic expression is characteristic of NPM1::ALK, which is the most common fusion. 
Cytotoxic molecules: TIA-1, perforin, granzyme B (IHC) Identification of a nonactivated (TIA1+ alone) or activated (perforin+ or granzyme B+) cytotoxic phenotype, characteristically seen in ALCL and a subset of PTCL-NOS. Many reactive cytotoxic cells may be present in noncytotoxic TCL (eg, TFHL-AI). 
TFH cell markers: PD1, ICOS, CXCL13, CD10, BCL6 (most commonly used), CD57, CXCR5, SAP, MAF, CD200
(IHC, FC for cell surface markers) 
TFH immunophenotype is defined by the expression of at least 2 and ideally 3 TFH markers in CD4+ cells. None of the TFH markers are in isolation sensitive or specific for the TFH phenotype.
Intensity of expression should be similar to that of reactive TFH cells, in a significant proportion of the presumed neoplastic cells. 
EBV (ISH, IHC) Detection mandatory to diagnose EBV-associated lymphomas.
Detection of EBV+ bystander B cells in TFH lymphomas, PTCL-NOS, and ATLL. 
EBER in situ hybridization is the gold standard for EBV detection; LMP-1 immunohistochemistry is less sensitive. 
CD56 (ICH, FC) Cytotoxic and NK cell lymphomas. Characteristically expressed in ENKTCL and MEITL; sometimes positive in primary nodal EBV+ PTCL; occasional aberrant expression in other T-cell lymphomas. 
Other markers of T helper cell subsets: TBX21, GATA3, CXCR3, CCR4, FOXP3 (ICH, FC for cell surface markers) Subsets of TH1 (TBX21+) and TH2 (GATA3+) PTCL-NOS.
FOXP3 (Treg) expression in a subset of ATLLs. 
Utility of these markers such as GATA3 or TBX21 is limited to specific diagnoses. Broad range of expression of GATA3 in nonhematologic malignancies and other lymphomas, including ALCLs. 
CD25 (ICH, FC) α-chain of the IL-2 receptor, expressed on normal T regulatory cells and usually expressed at high levels in ATLL cells, but also in ALCL and other PTCLs.  
Follicular dendritic cell (FDC) markers: CD21, CD23
(ICH) 
Expansion of FDC characteristic of TFHL-AI; demonstration of follicular pattern in TFHL-follicular. TFHL-AI pattern I has no FDC expansion. 
B-cell markers: CD20, CD79a, PAX5 (IHC, FC) Identification of a B-cell component or microenvironment in TFHL.
Abundant B cells in association with a T-cell lymphoproliferation favor a reactive over a malignant process. 
Some PTCLs may coexpress CD20 and/or other B-cell antigens.
PAX5 positivity in a subset of ALCLs. 
CD138, κ, λ (IHC) Plasma cells.
May be abundant in TFHL. 
Monotypic or even monoclonal plasma cells in some TFHLs. 
Antigen receptor gene rearrangements 
TRB and TRG (PCR- or NGS-based assays) Monoclonal gene rearrangements are detected in most T-cell neoplasms.
In some circumstances, lack of monoclonal TR gene rearrangement favors NK over T-cell derivation.
Monoclonal results support a diagnosis of TCL in cases with minimal involvement or when morphology and immunophenotyping are not definitively conclusive. 
Useful to demonstrate T-cell lineage in T-cell lymphomas with extensive antigen loss.
No correlation with TCRαβ+ vs TCRγδ+ phenotype.
Monoclonal TRB or TRG rearrangements may be detected in reactive T-cell lymphoproliferations (eg, viral infections), but are usually small. 
IGH, IGK, IGL (PCR- or NGS-based assays) Monoclonal gene rearrangements in general indicative of a B-cell neoplasm. Monoclonal IGH or IGK rearrangements may be detected in PTCLs with a B-cell component (especially TFHL). 
Specific genetic alterations 
ALK rearrangements (FISH) 100% of ALCL ALK+Immunohistochemistry for ALK usually used as a surrogate for genetic testing.
Rare cases of primary cutaneous ALCLs localized to the skin harbor ALK rearrangement and behave similarly to other ALK primary cutaneous cases. 
DUSP22 rearrangements (FISH) Subset (25%-30%) of ALCL ALKAlso in a subset of primary cutaneous ALCLs (and rare cases of lymphomatoid papulosis). 
TP63 rearrangements (FISH) Small subset (<10%) of ALCL ALK, and of PTCL-NOS. Expression of TP63 does not reliably distinguish cases with TP63 rearrangement (TP63 IHC+) vs those without TP63 rearrangement (may also be TP63 IHC+). 
JAK2 fusions (FISH or NGS) Subset of ALCL ALK, recurrent JAK2::STAT3 fusion in indolent clonal T-cell LPD of the gastrointestinal tract.  
TYK2 fusions (FISH or NGS) Subset of ALCL ALK, lymphomatoid papulosis and primary cutaneous ALCL. Various fusions reported, including NFκB2::TYK2, PABPC4::TYK2 NPM1::TYK2. 
CD28::ICOS and CD28::CTLA4 fusions (NGS) Subset of TFHL, rare in PTCL-NOS, cutaneous lymphomas, and ATLL.  
RHOAG17V (NGS, AS-PCR, ddPCR) Hot spot mutation in TFHL. Other RHOA variants occasional in TFH lymphomas and frequent in ATLL. 
IDH2 (NGS, AS-PCR, IHC) Hotspot mutations at R172 residue in one-third of TFHL-AI. Expression of mutant proteins can be demonstrated by IHC with antibodies specific of IDH2 variants.
Correlation with clear cell morphology. 
TET2, DNMT3 (NGS) Inactivating mutations, often multiple, very frequent in TFHL and less common in other PTCLs. Mutations also associated to clonal hematopoiesis and therefore not necessarily indicative of a T-cell neoplasm. 
CD28, PLCG1, FYN, CARD11, VAV1,… (NGS) Gain-of-function mutations recurrent in TFHL, ATLL, and cutaneous T-cell lymphomas, and rarely in other TCL.  
SETD2 (NGS and FISH) Loss-of-function mutations and/or deletions characteristic of MEITL and HSTL. Loss of SETD2 function translates to loss of H3K36me3 by IHC. 
JAK1, JAK3, STAT3, STAT5B (NGS) Gain-of-function mutations in various PTCL entities, especially frequent in primary intestinal TCLs, HSTL, ENKTCL, ALCL ALK, BIA-ALCL, leukemic NK-cell, and T-cell disorders.  
Iso(7q) (FISH) Iso(7q) detected in >80% of HSTL. Characteristic but not specific of HSTL, also reported in acute myeloid and lymphoblastic leukemia, myelodysplastic syndrome, rare cases of ENKTCL, and ALCL. 
MarkerDiagnostic utilityInterpretation and comments
Antigens assessed by immunohistochemistry (IHC) or flow cytometry (FC) 
Pan-T-cell antigens:
CD2, CD3, CD5, CD7, CD43 (IHC, FC) 
Identification T-cell (or NK) populations.
Aberrant loss of T-cell antigen expression is a phenotypic marker favoring neoplastic over benign lymphoproliferation. 
NK and T cells are distinguished by the expression of surface CD3 (assessed on fresh cells by FC), positive in T cells and negative in NK cells, whereas both T and NK cells express cytoplasmic CD3 (assessed by IHC on routinely processed tissues); in addition, CD5 is constitutively absent in NK cells (and in TCRγδ+ T cells)
Extensive loss of T-cell antigens and possibly «null» immunophenotype is characteristic of ALK+ and to a lesser extent ALK ALCL.
CD7 loss is typical of ATLL. 
CD4 and CD8 (IHC, FC) Characterization of T-cell proliferations according to the CD4+CD8 or CD4CD8+ lineages. Double negative expression may indicate immature lymphoid cells, TCRγδ+ or NK cells, or aberrant phenotype in mature TCRαβ+ T cells, found in ALPS or lymphomas.
Double positive expression may indicate immature lymphoid cells, or aberrant phenotype in mature TCRαβ+ T cells, usually indicative of malignancy. 
TCRβ and TCRδ chains (IHC, FC) Identification of TCRαβ+ vs TCRγδ+ T cells. Most nodal PTCLs derive from TCRαβ+ T cells.
PTCL may show TCR downregulation, or silencing of TCR expression. 
TRBC1 (IHC, FC) Identification of clonal T-cell population by monotypic expression pattern. Only applicable if surface CD3 and TCRαβ expression is retained. 
CD30 (ICH, FC) Activation marker, not lineage-specific.
Strong uniform expression in ALCLs, heterogeneous expression in a proportion of cases in many TCL entities, usually in a subset of the cells. 
Only scattered cells may be positive in small-cell and histiocyte-rich patterns of ALK+ ALCL.
CD30 expression in bystander Reed–Sternberg-like cells in TFHL and PTCL-NOS.
May be targetable by antibody–drug conjugates. 
CD15 (IHC, FC) Glycan determinant, marker of myeloid cells.
May be positive in the neoplastic cells in ALCL and a subset of PTCL-NOS. 
Coexpression of CD30 and CD15 in bystander HRS B cells in various PTCLs. 
ALK (IHC) Demonstration of ALK expression is essential for the diagnosis of ALK+ ALCL. IHC is an excellent surrogate to ALK gene fusion detection by genetic assays. Variable patterns of subcellular staining according to the partner gene of ALK in the fusion.
Nuclear and cytoplasmic expression is characteristic of NPM1::ALK, which is the most common fusion. 
Cytotoxic molecules: TIA-1, perforin, granzyme B (IHC) Identification of a nonactivated (TIA1+ alone) or activated (perforin+ or granzyme B+) cytotoxic phenotype, characteristically seen in ALCL and a subset of PTCL-NOS. Many reactive cytotoxic cells may be present in noncytotoxic TCL (eg, TFHL-AI). 
TFH cell markers: PD1, ICOS, CXCL13, CD10, BCL6 (most commonly used), CD57, CXCR5, SAP, MAF, CD200
(IHC, FC for cell surface markers) 
TFH immunophenotype is defined by the expression of at least 2 and ideally 3 TFH markers in CD4+ cells. None of the TFH markers are in isolation sensitive or specific for the TFH phenotype.
Intensity of expression should be similar to that of reactive TFH cells, in a significant proportion of the presumed neoplastic cells. 
EBV (ISH, IHC) Detection mandatory to diagnose EBV-associated lymphomas.
Detection of EBV+ bystander B cells in TFH lymphomas, PTCL-NOS, and ATLL. 
EBER in situ hybridization is the gold standard for EBV detection; LMP-1 immunohistochemistry is less sensitive. 
CD56 (ICH, FC) Cytotoxic and NK cell lymphomas. Characteristically expressed in ENKTCL and MEITL; sometimes positive in primary nodal EBV+ PTCL; occasional aberrant expression in other T-cell lymphomas. 
Other markers of T helper cell subsets: TBX21, GATA3, CXCR3, CCR4, FOXP3 (ICH, FC for cell surface markers) Subsets of TH1 (TBX21+) and TH2 (GATA3+) PTCL-NOS.
FOXP3 (Treg) expression in a subset of ATLLs. 
Utility of these markers such as GATA3 or TBX21 is limited to specific diagnoses. Broad range of expression of GATA3 in nonhematologic malignancies and other lymphomas, including ALCLs. 
CD25 (ICH, FC) α-chain of the IL-2 receptor, expressed on normal T regulatory cells and usually expressed at high levels in ATLL cells, but also in ALCL and other PTCLs.  
Follicular dendritic cell (FDC) markers: CD21, CD23
(ICH) 
Expansion of FDC characteristic of TFHL-AI; demonstration of follicular pattern in TFHL-follicular. TFHL-AI pattern I has no FDC expansion. 
B-cell markers: CD20, CD79a, PAX5 (IHC, FC) Identification of a B-cell component or microenvironment in TFHL.
Abundant B cells in association with a T-cell lymphoproliferation favor a reactive over a malignant process. 
Some PTCLs may coexpress CD20 and/or other B-cell antigens.
PAX5 positivity in a subset of ALCLs. 
CD138, κ, λ (IHC) Plasma cells.
May be abundant in TFHL. 
Monotypic or even monoclonal plasma cells in some TFHLs. 
Antigen receptor gene rearrangements 
TRB and TRG (PCR- or NGS-based assays) Monoclonal gene rearrangements are detected in most T-cell neoplasms.
In some circumstances, lack of monoclonal TR gene rearrangement favors NK over T-cell derivation.
Monoclonal results support a diagnosis of TCL in cases with minimal involvement or when morphology and immunophenotyping are not definitively conclusive. 
Useful to demonstrate T-cell lineage in T-cell lymphomas with extensive antigen loss.
No correlation with TCRαβ+ vs TCRγδ+ phenotype.
Monoclonal TRB or TRG rearrangements may be detected in reactive T-cell lymphoproliferations (eg, viral infections), but are usually small. 
IGH, IGK, IGL (PCR- or NGS-based assays) Monoclonal gene rearrangements in general indicative of a B-cell neoplasm. Monoclonal IGH or IGK rearrangements may be detected in PTCLs with a B-cell component (especially TFHL). 
Specific genetic alterations 
ALK rearrangements (FISH) 100% of ALCL ALK+Immunohistochemistry for ALK usually used as a surrogate for genetic testing.
Rare cases of primary cutaneous ALCLs localized to the skin harbor ALK rearrangement and behave similarly to other ALK primary cutaneous cases. 
DUSP22 rearrangements (FISH) Subset (25%-30%) of ALCL ALKAlso in a subset of primary cutaneous ALCLs (and rare cases of lymphomatoid papulosis). 
TP63 rearrangements (FISH) Small subset (<10%) of ALCL ALK, and of PTCL-NOS. Expression of TP63 does not reliably distinguish cases with TP63 rearrangement (TP63 IHC+) vs those without TP63 rearrangement (may also be TP63 IHC+). 
JAK2 fusions (FISH or NGS) Subset of ALCL ALK, recurrent JAK2::STAT3 fusion in indolent clonal T-cell LPD of the gastrointestinal tract.  
TYK2 fusions (FISH or NGS) Subset of ALCL ALK, lymphomatoid papulosis and primary cutaneous ALCL. Various fusions reported, including NFκB2::TYK2, PABPC4::TYK2 NPM1::TYK2. 
CD28::ICOS and CD28::CTLA4 fusions (NGS) Subset of TFHL, rare in PTCL-NOS, cutaneous lymphomas, and ATLL.  
RHOAG17V (NGS, AS-PCR, ddPCR) Hot spot mutation in TFHL. Other RHOA variants occasional in TFH lymphomas and frequent in ATLL. 
IDH2 (NGS, AS-PCR, IHC) Hotspot mutations at R172 residue in one-third of TFHL-AI. Expression of mutant proteins can be demonstrated by IHC with antibodies specific of IDH2 variants.
Correlation with clear cell morphology. 
TET2, DNMT3 (NGS) Inactivating mutations, often multiple, very frequent in TFHL and less common in other PTCLs. Mutations also associated to clonal hematopoiesis and therefore not necessarily indicative of a T-cell neoplasm. 
CD28, PLCG1, FYN, CARD11, VAV1,… (NGS) Gain-of-function mutations recurrent in TFHL, ATLL, and cutaneous T-cell lymphomas, and rarely in other TCL.  
SETD2 (NGS and FISH) Loss-of-function mutations and/or deletions characteristic of MEITL and HSTL. Loss of SETD2 function translates to loss of H3K36me3 by IHC. 
JAK1, JAK3, STAT3, STAT5B (NGS) Gain-of-function mutations in various PTCL entities, especially frequent in primary intestinal TCLs, HSTL, ENKTCL, ALCL ALK, BIA-ALCL, leukemic NK-cell, and T-cell disorders.  
Iso(7q) (FISH) Iso(7q) detected in >80% of HSTL. Characteristic but not specific of HSTL, also reported in acute myeloid and lymphoblastic leukemia, myelodysplastic syndrome, rare cases of ENKTCL, and ALCL. 

AI, angioimmunoblastic type; ALCL, anaplastic large cell lymphoma; ALK, anaplastic lymphoma kinase; ALPS, autoimmune lymphoproliferative syndrome; AS-PCR, allele-specific PCR; ATLL, adult T-cell leukemia/lymphoma; BIA, breast implant–associated; ddPCR, digital droplet PCR; EBERs, EBV-encoded small RNAs; EBV, Epstein-Barr virus; ENKTCL, extranodal NK/T-cell lymphoma, nasal type; F, follicular; FC, flow cytometry; FDC, follicular dendritic cell; FISH, fluorescent in situ hybridization; HSTL, hepatosplenic T-cell lymphoma; IHC, immunohistochemistry; IL-2, interleukin-2; LPD, lymphoproliferative disorder; LMP-1, latent membrane protein 1; MEITL, monomorphic epitheliotropic intestinal T-cell lymphoma; NGS, high-throughput sequencing; PCR, polymerase chain reaction; PTCL-NOS, peripheral TCL, not otherwise specified; TCL, T-cell lymphoma; TFHL, follicular helper TCL.

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