Table 2.

Germ line mutations and acquired conditions predisposing to development of NK/T-cell lymphomas and leukemias

Predisposing featureLymphomasComments
Germ line mutations 
Ataxia-telangiectasia T-lymphoblastic leukemia/lymphoma in children
T-cell prolymphocytic leukemia (T-PLL) in young adults 
Autosomal recessive disease with biallelic loss-of-function ATM mutations. Monoallelic mutations predispose to T-PLL in older individuals. 
Nijmegen breakage syndrome T-lymphoblastic leukemia/lymphoma Autosomal recessive disease with biallelic NBS mutations. 
HAVCR2 mutations9  Subcutaneous panniculitis-like T-cell lymphoma (SPTCL) 25%-60% of patients with SPTCL harbor germ line HAVCR2 mutations and more often present at a younger age with hemophagocytic lymphohistiocytosis. 
TET2 mutations10,11  Mature T-cell lymphoma reported in some cases (follicular helper T-cell lymphoma, ALK anaplastic large cell lymphoma [ALCL]) in childhood or adulthood Usually biallelic homozygous or heterozygous germ line mutations cause hypermethylation of peripheral blood DNA, childhood immunodeficiency with autoimmune lymphoproliferative syndrome (ALPS)-like features, and secondary development of various lymphomas. 
Acquired conditions 
HTLV-1 infection Adult T-cell leukemia/lymphoma Often by transmission of HTLV-1 through breastfeeding. 
Chronic active EBV disease Extranodal NK/T-cell lymphoma, nasal type, or aggressive NK-cell leukemia Risk of progression to lymphoma or leukemia in patients with the systemic type of chronic active EBV disease not treated with allogeneic bone marrow transplantation. 
Breast implant12,13  Breast implant–associated ALCL (BIA-ALCL) Textured breast implants are a risk factor for the development of BIA-ALCL, and the risk is increased in women with genetic predisposition to breast cancer (BRCA1 or TP53 germ line mutations). 
Celiac disease Refractory celiac disease, type II (RCD-II) and enteropathy-associated T-cell lymphoma (EATL) RCD II/EATL is a rare complication of celiac disease, an immune-mediated enteropathy occurring in genetically susceptible individuals carrying HLA-DQ2 or HLA-DQ8.
1% of celiac disease patients will develop RCD-II. The 5-y risk of progression to EATL for individuals with RCD-II is 30%-50%. 
Immune suppression Hepatosplenic T-cell lymphoma (HSTL) Many patients with HSTL have preexisting iatrogenic immune deregulation to manage solid organ transplantation, inflammatory bowel disease, or rheumatoid arthritis, and often received anti-TNFalpha or other immunomodulator agents. Given the rarity of HSTL, the risk is not quantifiable. 
Chronic lymphocytic leukemia (CLL)14  Cutaneous T-cell lymphoma and systemic lymphoma (peripheral T-cell lymphoma, NOS, ALCLs, follicular helper T-cell lymphoma) The increased risk for developing cutaneous or systemic PTCLs in patients with CLL has been linked to the accumulation of oligo- or monoclonal T-cell populations with abnormal phenotypes. 
Clonal hematopoiesis (CH)15,16  Follicular helper T-cell lymphoma (TFHL) Most patients with TFHL have underlying clonally linked CH often with mutations in TET2 and DNMT3A. Data suggest that a subset of peripheral T-cell lymphoma, NOS are also related to CH. 
Lymphocytic variant eosinophilic syndrome (LHES)17  Peripheral T-cell lymphoma, NOS, follicular helper T-cell lymphoma LHES is a form of chronic indolent T-cell LPD encompassing a clonal and phenotypically aberrant (usually sCD3 CD4+) T-cell population in blood. A subset of the patients (up to 25%) may progress to lymphoma. 
T-cell clones of uncertain significance (T-CUS)18  T-cell large granular lymphocytic leukemia (T-LGLL) Premalignant condition characterized by the presence of small clonal cell expansions in individuals without symptoms or signs of T-LGLL. 
Predisposing featureLymphomasComments
Germ line mutations 
Ataxia-telangiectasia T-lymphoblastic leukemia/lymphoma in children
T-cell prolymphocytic leukemia (T-PLL) in young adults 
Autosomal recessive disease with biallelic loss-of-function ATM mutations. Monoallelic mutations predispose to T-PLL in older individuals. 
Nijmegen breakage syndrome T-lymphoblastic leukemia/lymphoma Autosomal recessive disease with biallelic NBS mutations. 
HAVCR2 mutations9  Subcutaneous panniculitis-like T-cell lymphoma (SPTCL) 25%-60% of patients with SPTCL harbor germ line HAVCR2 mutations and more often present at a younger age with hemophagocytic lymphohistiocytosis. 
TET2 mutations10,11  Mature T-cell lymphoma reported in some cases (follicular helper T-cell lymphoma, ALK anaplastic large cell lymphoma [ALCL]) in childhood or adulthood Usually biallelic homozygous or heterozygous germ line mutations cause hypermethylation of peripheral blood DNA, childhood immunodeficiency with autoimmune lymphoproliferative syndrome (ALPS)-like features, and secondary development of various lymphomas. 
Acquired conditions 
HTLV-1 infection Adult T-cell leukemia/lymphoma Often by transmission of HTLV-1 through breastfeeding. 
Chronic active EBV disease Extranodal NK/T-cell lymphoma, nasal type, or aggressive NK-cell leukemia Risk of progression to lymphoma or leukemia in patients with the systemic type of chronic active EBV disease not treated with allogeneic bone marrow transplantation. 
Breast implant12,13  Breast implant–associated ALCL (BIA-ALCL) Textured breast implants are a risk factor for the development of BIA-ALCL, and the risk is increased in women with genetic predisposition to breast cancer (BRCA1 or TP53 germ line mutations). 
Celiac disease Refractory celiac disease, type II (RCD-II) and enteropathy-associated T-cell lymphoma (EATL) RCD II/EATL is a rare complication of celiac disease, an immune-mediated enteropathy occurring in genetically susceptible individuals carrying HLA-DQ2 or HLA-DQ8.
1% of celiac disease patients will develop RCD-II. The 5-y risk of progression to EATL for individuals with RCD-II is 30%-50%. 
Immune suppression Hepatosplenic T-cell lymphoma (HSTL) Many patients with HSTL have preexisting iatrogenic immune deregulation to manage solid organ transplantation, inflammatory bowel disease, or rheumatoid arthritis, and often received anti-TNFalpha or other immunomodulator agents. Given the rarity of HSTL, the risk is not quantifiable. 
Chronic lymphocytic leukemia (CLL)14  Cutaneous T-cell lymphoma and systemic lymphoma (peripheral T-cell lymphoma, NOS, ALCLs, follicular helper T-cell lymphoma) The increased risk for developing cutaneous or systemic PTCLs in patients with CLL has been linked to the accumulation of oligo- or monoclonal T-cell populations with abnormal phenotypes. 
Clonal hematopoiesis (CH)15,16  Follicular helper T-cell lymphoma (TFHL) Most patients with TFHL have underlying clonally linked CH often with mutations in TET2 and DNMT3A. Data suggest that a subset of peripheral T-cell lymphoma, NOS are also related to CH. 
Lymphocytic variant eosinophilic syndrome (LHES)17  Peripheral T-cell lymphoma, NOS, follicular helper T-cell lymphoma LHES is a form of chronic indolent T-cell LPD encompassing a clonal and phenotypically aberrant (usually sCD3 CD4+) T-cell population in blood. A subset of the patients (up to 25%) may progress to lymphoma. 
T-cell clones of uncertain significance (T-CUS)18  T-cell large granular lymphocytic leukemia (T-LGLL) Premalignant condition characterized by the presence of small clonal cell expansions in individuals without symptoms or signs of T-LGLL. 

EBV, Epstein-Barr virus; HTLV-1, human T-lymphotropic virus 1; TNF, tumor necrosis factor.

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